Ladon was the serpent-like dragon that twined and twisted around the tree in the Garden of the Hesperides and guarded the golden apples. He was overcome by Heracles by a bow that he had. The following day, Jason and the Argonauts passed by on their chthonic return journey from Colchis and heard the lament of “shining” Aegle, one of the four Hesperides, and viewed the still-twitching Ladon.